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56

"Tsee tsee tsee tseeeeeeeeeeeeeee
tseeeeeeeeeeeeee tseeeeeeeeeeee." Then finally see animals. Lots of fusucollis. Jumping about very excitedly high in rather tall trees, 30-70 ft. up. Continue vocalizing as before for some seconds. Now it is obvious that they are uttering ordinary LSN's and Whistles of the species. Whistles still greatly predominant. Then the animals become aware of our presence. Usual alarm reactions. Tail curving LSN's.  Trill's. Whistles almost stop. At same time; it becomes evident that animals are separated into two groups. Approx. 10-20 ft apart. One group consists of 2 inds. The other includes 4 or 5. The animals gradually become habituated. Stop uttering alarm calls. But do not resume Whistling. Just sit. Group of 4 or 5 largely hidden. Group of 2 in plain view. Both inds. of this last raise their tails forward and upward, and then scratch them vigorously with their hands. Displacement??? Then the animals move off. I am fairly certain that the two groups go in different directions. 

[[sketch - two arrows pointing in opposite directions]]

This must have been a real boundary dispute. (And of course, one might expect face-to-face territorial disputes to be more common in this species than in geoffroyi. Simply because fuscicollis lacks the long list once repellent 2W pattern.) 

This incident would also indicate that the Whistle of fuscicollis is primarily an intra-specific signal. Which, in turn, probably indicates that it contains relatively more attack than either the LSN or the Trill. 

Possibly some sequences of LSN's-Whistles is 

Transcription Notes:
.inds. = individuals last syllable on page carried to beginning of next page to complete word per Smithsonian